An African grey parrot spontaneously showed he understood the concept of zero

Alex, a parrot trained for decades by researcher Irene Pepperberg, had learned to name quantities and objects but was never taught the word 'zero.' During a test, shown a tray with nothing matching the question, he spontaneously answered 'none' — a label he'd only ever used before to describe an absence of similarity between two objects. It made him the first bird shown to grasp something like the concept of zero.

— Irene Pepperberg and Jesse Gordon, Number Comprehension by a Grey Parrot (Psittacus erithacus), Including a Zero-Like Concept — Journal of Comparative Psychology, 2005

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